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Will removing the string tree cause any issues with the ebanol nut?
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string trees are there to help give a better break angle from the nut to the tuners, they are added to necks that have a straight headstock. The 540S has a headstock that bends backwards so no string trees are needed.
As far as what locking tuners will fit... gotoh makes some auto locking tuners that dont require any extra holes to be drilled, they should fit just like the original tuners.
I recently put some sperzel locking tuners in my rt150 because they offer "graduated height" tuners which allow you to remove the string tree because the tuners go from higher string height to lower string height on the headstock.
the sperzels and some others will require you to drill a small hole on the backside of the headstock to install them. its really easy but it will leave the original hole behind.