When you strum a guitar hard, there is more noticeable vibration taking place at the center of the string (which is closer to the neck pickup position) than on either end. That makes it much easier for a pickup to do its job. As a result, a much less senstive pickup is required at the
neck position and a much more senstitive pickup is required at the bridge (hence the larger DC resistance readings for
bridge pickups).
I would guess the the reduction of vibration at the end of the string would attenuate the low frequencies more than the high frequencies, resulting in a "mellower" or "darker" neck tone and a "more edgy" and "brighter" bridge tone.
Of course, the pickups can be voiced to compensate for these natural tendencies. For example, the Humbucker from Hell provides a very bright neck tone and the Tone Zone provides a darker and more bassy bridge tone.