The 521 is the loss leader of all the fixed just as the 550's are of the trems. The extra gets you Dimarzio's, the Prestige case, and a multi tool which doesn't add up to $400, but it's not supposed to.
All Prestige fret end finishing is a bit lame currently. I just checked in 2 521's and they were a hair lamer than the RGR I just looked at.